National Football League players sat out, knelt and linked arms during pre-game national anthems played across the country and in London today, hours after US President Donald Trump called on fans to boycott teams that do not discipline players who protest.

In the first few games since Trump stepped up his criticism of NFL players, dozens of players and coaches of teams including the Baltimore Ravens, Jacksonville Jaguars, Philadelphia Eagles and Miami Dolphins did not stand for the anthem and took a knee, a gesture that began last year as a protest over police treatment of African-Americans and other minorities.

The Pittsburgh Steelers waited off the field during the national anthem before their game against the Chicago Bears in Chicago today afternoon to avoid "playing politics" in divisive times, coach Mike Tomlin said.

And in Philadelphia, city police officers joined with Eagles players and team owner Jeffrey Lurie to link arms during the anthem in a sign of solidarity.

The defiant moves came in response to a series of comments made by Trump over the weekend criticizing players who refuse to stand for the national anthem.

Jacksonville Jaguars players kneel during the national anthem. Photo: ReutersJacksonville Jaguars players kneel during the national anthem. Photo: Reuters

"If NFL fans refuse to go to games until players stop disrespecting our Flag & Country, you will see change take place fast," Trump wrote this morning on Twitter. "Fire or suspend!"

In a second tweet, Trump, who is spending the weekend at his golf club in Bedminster, New Jersey, said that the "league should back" fans who are upset about the protests.

The form of protest began in 2016 when then San Francisco 49ers quarterback Colin Kaepernick put one knee to the ground during pre-game renditions of the "Star Spangled Banner." Several players have since made similar gestures of protest before games.

Kaepernick initially started his protests last year by sitting during the anthem but when critics said it was a sign of disrespect, he instead took to kneeling.

At a political rally in Alabama on Friday, Trump suggested any protesting football player was a "son of a bitch" and should lose his job.

This morning, NFL managers, coaches and owners began to weigh in on the escalating feud, criticizing the president for calling out players' political dissent.

New England Patriots Chairman and CEO Robert Kraft, who has dined with Trump and who the president considers a friend, criticised Trump and defended players' right to protest.

Members of the Indianapolis Colts kneel ahead of their game against the Cleveland Browns. Photo: ReutersMembers of the Indianapolis Colts kneel ahead of their game against the Cleveland Browns. Photo: Reuters

"I am deeply disappointed by the tone of the comments made by the president on Friday," Kraft said in a statement posted on Twitter. Kraft said he supports his players' "right to peacefully affect social change and raise awareness in a manner they feel is most impactful."

Jaguars owner Shad Khan linked arms with team players in solidarity at the game against the Ravens in London's Wembley Stadium. Khan donated $1 million to the Trump inauguration fund.

And Robert McNair, chairman of the Houston Texans team, said that Trump's comments were "divisive and counterproductive to what our country needs right now." McNair also donated $1 million to Trump.

NFL Commissioner Roger Goodell said Trump's comments in Alabama revealed an "unfortunate lack of respect" for the NFL and its players.

As commissioner, Goodell reports to NFL owners, some of whom have supported Trump. New York Jets owner Woody Johnson, a major Trump presidential campaign donor, was confirmed by the Senate last month as Trump's pick to serve as US ambassador to Britain.

The union representing professional football players also rejected Trump's comments, saying it would defend their right to freedom of expression.

In Chicago, Trump's feud was the main topic of conversation at the South Loop Club sports bar.

"This is a first amendment issue and the president is supposed to uphold that right," said Sam Cunningham, 55, who was watching the game with his wife. "He should know better than anyone that to fire someone because of their opinion is not right."

National Basketball Association players also struck back against comments by the president yesterday after Trump clashed with one of the biggest stars in the NBA, Stephen Curry. Then on Saturday evening, Oakland As rookie Bruce Maxwell became the first Major League Baseball player to kneel for the national anthem in protest.

In a Twitter message yesterday, the president rescinded a White House invitation to Stephen Curry, who had said he would "vote" against the planned visit by the NBA champion Golden State Warriors.

The protest has infuriated president Trump. Photo: ReutersThe protest has infuriated president Trump. Photo: Reuters

Curry told a news conference in Oakland, California: "It's beneath the leader of a country to go that route."

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